A fertility clinic affiliated with the University of Connecticut Health Center mistakenly implanted one woman with another woman’s fertilized embryo in 2009, but the doctor who runs the clinic still earned more than $800,000 last year making him the fifth highest-paid state employee.

Dr. John Nulsen is the medical director of the Center for Advanced Reproductive Services where he received $881,472.44 last year. Only the dean of the health center and UConn’s football and basketball coaches made more.

In 2009, the year of the mistake, Nulsen received $796,808, including a bonus of $537,720. He earned $889,784 in 2008 and $662,034 in 2007, according to CTSunlight.org.

CARS, the private clinic, by contract reimburses the health center for Nulsen’s bonuses, according to a health center spokesman.

The state has already paid a $45,000 settlement related to the embryo mix-up and a $2 million claim against the state is pending.